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Volvocaceae
The Volvocaceae are a family of unicellular or colonial biflagellates, including the typical genus ''Volvox''. The family was named by Ehrenberg in 1834, and is known in older classifications as the Volvocidae. All species are colonial and inhabit freshwater environments.
==Description==
The simplest of the Volvocaeans are ordered assemblies of cells, each similar to the related unicellar protist ''Chlamydomonas'' and embedded in a gelatinous matrix. In the genus ''Gonium'', for example, each individual organism is a flat plate consisting of 4 to 16 separate cells, each with two flagella. Similarly, the genera ''Eudorina'' and ''Pandorina'' form hollow spheres, the former consisting of 16 cells, the latter of 32 to 64 cells. In these genera each cell can reproduce a new organism by mitosis.〔Scott 2000〕
Other genera of Volvocaceans represent another principle of biological development as each organism develops differented cell types. In ''Pleodorina'' and ''Volvox'', most cells are somatic and only a few are reproductive. In ''Pleodorina californica'' a colony normally has either 128 or 64 cells, of which those in the anterior region have only a somatic function, while those in the posterior region can reproduce; the ratio being 3:5. In ''Volvox'' only very few cells are able to reproduce new individuals, and in some species of ''Volvox'' the reproductive cells are derived from cells looking and behaving like somatic cells. In ''V. carteri'', on the other hand, the division of labor is complete with reproductive cells being set aside during cell division, and they never assume somatic functions or develop functional flagella.〔
Thus, the simplest Volvocaceans are colonial organisms but others are truly multicellular organisms. Larger volvocaceans have evolved a specialized form of heterogamy called oogamy, the production of small motile sperm by one mating type and relatively larger immotile eggs by another. Among the Volvocaceans are thus the simplest organisms with distinguishable male and female members. In all Volvocaceans, the fertilization reaction results in the production of a dormant diploid zygote (zygospore) capable of surviving in harsh environments. Once conditions have improved the zygospore germinates and undergoes meiosis to produce haploid offspring of both mating types.〔
抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』
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