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Roupala : ウィキペディア英語版
Roupala

''Roupala'' is a neotropical genus of woody shrubs and trees in the plant family Proteaceae. Its 34 species are generally found in forests from sea level to 4000 m altitude from Mexico to Argentina.
==Taxonomy and naming==
The genus was defined by Jean Baptiste Christophore Fusée Aublet in 1775, its name derived from a local name ''roupale'' in French Guiana.〔
In their 1975 monograph on the Proteaceae, Lawrie Johnson and Barbara Briggs placed it in a subtribe Roupalinae alongside the New Caledonian genus ''Kermadecia'' as the genera had similar floral parts and leaves. Both taxa also have 14 chromosome pairs.
In 2006, the family's classification was redefined using molecular data. Here, ''Roupala'' emerged as a sister to the genera ''Orites'' and ''Neorites'', with ''Knightia'' as the next most closely related taxon, while ''Kermadecia'' was not related. They thus placed the first three genera in the subtribe ''Roupalinae'', conceding that the next closest relatives of this group is unclear. This group lies within the subfamily Grevilleoideae.
The genus likely originated in Gondwana before South America split away around 110 million years ago, and then spread into Central America in the Miocene around six million years ago when the Americas came into contact with each other. Clock dating with molecular and fossil data indicated ancestors of the genus may have split from ''Neorites'' in the mid-Oligocene around 30 million years ago, and that this lineage in turn separated from the ancestors of ''Orites'' in the late Eocene around 36 million years ago.

抄文引用元・出典: フリー百科事典『 ウィキペディア(Wikipedia)
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