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Monticolomys : ウィキペディア英語版
Monticolomys

''Monticolomys'' is a genus of rodents within the subfamily Nesomyinae of the family Nesomyidae, and is closely related to ''Macrotarsomys''. The only species, ''Monticolomys koopmani'', also known as the Malagasy mountain mouse or Koopman's montane voalavo, is found in the highlands of eastern Madagascar. A small mouse-like rodent, ''M. koopmani'' is dark brown on the upperparts and dark gray below. It has small, rounded, densely haired ears and broad feet with well-developed pads. The long tail lacks a tuft at the tip. The skull is delicate and lacks crests and ridges on its roof.
First collected in 1929, ''Monticolomys koopmani'' was not formally described until 1996, but it is now known to have a broad distribution. Active during the night, it occurs in both montane forest and human-disturbed grasslands and feeds on fruits and seeds. A scansorial animal, it climbs trees but also lives on the ground. Although habitat destruction may pose a threat, it is classified as "Least Concern" on the IUCN Red List.
==Taxonomy==
A specimen of ''Monticolomys koopmani'' was captured in 1929 during the Mission Zoologique Franco-Anglo-Américaine to Madagascar, but the rodents obtained by the expedition were never studied in detail. It was not until the 1970s that Karl Koopman and Guy Musser recognized that the animal—whose skin had landed at the American Museum of Natural History in New York, while the skull was at the Muséum national d'histoire naturelle in Paris—represented an otherwise unknown species. However, they never published their results. In 1993, Steven Goodman rediscovered the species on Madagascar and in 1996 he and Michael Carleton finally published a formal description.〔Carleton and Goodman, 1996, p. 232〕 They named the animal ''Monticolomys koopmani'', as the sole member of a new genus.〔Carleton and Goodman, 1996, p. 233〕 The generic name ''Monticolomys'' means "mountain-dwelling mouse" and refers to the animal's montane habitat, and the specific name ''koopmani'' honors Karl Koopman for his many contributions to mammalian systematics.〔Carleton and Goodman, 1996, p. 242〕 Common names in use for the animal include "Koopman's Montane Voalavo"〔Musser and Carleton, 2005, p. 952〕 and "Malagasy Mountain Mouse".〔
The indigenous rodents of Madagascar, the Nesomyinae, prior to the discovery of ''Monticolomys'' comprised seven very distinctive genera—so distinct from each other that some have found it difficult to accept that they are closely related.〔Carleton and Goodman, 1996, p. 249〕 ''Monticolomys'', however, does not follow this pattern, in that it is similar and closely related to the gerbil-like genus ''Macrotarsomys'' of western Madagascar. This relationship was originally proposed by Goodman and Carleton based on morphology,〔Carleton and Goodman, 1996, p. 250〕 and was strongly supported by a DNA sequence analysis (based on the cytochrome ''b'' gene) published in 1999.〔Jansa et al., 1999, pp. 262–263〕 While this study provided some weak support for a relationship between the ''Macrotarsomys–Monticolomys'' clade and the giant jumping rat, ''Hypogeomys'',〔Jansa et al., 1999, p. 263〕 a later study based on the IRBP gene instead placed ''Macrotarsomys–Monticolomys'' sister to a clade containing four other nesomyine genera—''Eliurus'', ''Voalavo'', ''Gymnuromys'', and ''Brachytarsomys''.〔Jansa and Weksler, 2004, fig. 1〕

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