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ruminant : ウィキペディア英語版
ruminant

Ruminants are mammals that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microbial actions. The process typically requires the fermented ingesta (known as cud) to be regurgitated and chewed again. The process of rechewing the cud to further break down plant matter and stimulate digestion is called rumination. The word "ruminant" comes from the Latin ''ruminare'', which means "to chew over again".
The roughly 150 species of ruminants include both domestic and wild species. Ruminating mammals include cattle, goats, sheep, giraffes, yaks, deer, antelope, and some macropods.〔Fowler, M.E. (2010). "Medicine and Surgery of Camelids", Ames, Iowa: Wiley-Blackwell. Chapter 1 General Biology and Evolution addresses the fact that camelids (including camels and llamas) are not ruminants, pseudo-ruminants, or modified ruminants.〕
Taxonomically, the suborder Ruminantia (also known as ruminants) is a lineage of herbivorous artiodactyls that includes the most advanced and widespread of the world's ungulates. The term 'ruminant' is not synonymous with Ruminantia. Suborder Ruminantia includes many ruminant species, but does not include tylopods and marsupials.〔
==Explanation==

The primary difference between a ruminant and nonruminant is that ruminants have a four-compartment stomach. The four parts are the rumen, reticulum, omasum, and abomasum. In the first two chambers, the rumen and the reticulum, the food is mixed with saliva and separates into layers of solid and liquid material. Solids clump together to form the cud or bolus.
The cud is then regurgitated and chewed to completely mix it with saliva and to break down the particle size. Fiber, especially cellulose and hemicellulose, is primarily broken down in these chambers by microbes (mostly bacteria, as well as some protozoa, fungi and yeast) into the three volatile fatty acids (VFAs): acetic acid, propionic acid, and butyric acid. Protein and nonstructural carbohydrate (pectin, sugars, and starches) are also fermented.
Though the rumen and reticulum have different names, they represent the same functional space as digesta can move back and forth between them. Together, these chambers are called the reticulorumen. The degraded digesta, which is now in the lower liquid part of the reticulorumen, then passes into the next chamber, the omasum, where water and many of the inorganic mineral elements are absorbed into the blood stream.
After this, the digesta is moved to the true stomach, the abomasum. The abomasum is the direct equivalent of the monogastric stomach, and digesta is digested here in much the same way. Digesta is finally moved into the small intestine, where the digestion and absorption of nutrients occurs. Microbes produced in the reticulorumen are also digested in the small intestine. Fermentation continues in the large intestine in the same way as in the reticulorumen.
Only small amounts of glucose are absorbed from dietary carbohydrates. Most dietary carbohydrates are fermented into VFAs in the rumen. The glucose needed as energy for the brain and for lactose and milk fat in milk production, as well as other uses, comes from nonsugar sources, such as the VFA propionate, glycerol, lactate, and protein. The VFA propionate is used for around 70% of the glucose and glycogen produced and protein for another 20% (50% under starvation conditions).〔William O. Reece (2005). (Functional Anatomy and Physiology of Domestic Animals ), pages 357-358 ISBN 978-0-7817-4333-4〕〔Colorado State University, Hypertexts for Biomedical Science: (Nutrient Absorption and Utilization in Ruminants )〕

抄文引用元・出典: フリー百科事典『 ウィキペディア(Wikipedia)
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